Tron asks Anon to guard the ceremony but finds a suspicious female ISO, Quorra, who tries to talk her way past the guards, prompting Anon to follow her. Radia, leader of the ISOs, is at a formal ceremony to make her a System Administrator alongside Clu. Flynn has got Anon to try to control the system. Jalen, one of the leaders of the ISOs, recently died and Flynn suspects the murder was organized by Clu, a second version of the original program from the first film. The game starts with video footage of Kevin Flynn who is discussing the existence of ISOs (Isomorphic Algorithms), a group of programs with a measure of free will that has emerged spontaneously on the Grid, and are disliked by the Basic programs. The player controls a system monitor program named "Anon" (short for anonymous) - a security program owned by Flynn to investigate a conspiracy in the world of Tron. Tron: Evolution explains the events that led to Kevin Flynn's imprisonment inside the Grid, as well as telling how the Grid evolved through the years. It is designed to be an integral part of the Tron story, featuring certain characters and settings from both films.
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Tron: Evolution serves as a precursor to the movie Tron: Legacy and sequel to both the graphic novel Tron: Betrayal and the Nintendo Wii and DS versions, Tron: Evolution - Battle Grids.
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Five more maps and the Black Guard character skin were released in a downloadable pack one of these, the "Classic" map, is a replica of the light cycle grid from the original Tron film.
Two more multiplayer maps, as well as a Sam Flynn character skin, were made available at the game's launch as downloadable content through codes included in the game package.
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After becoming version 50, the players cannot upgrade own character anymore. The character can gain versions up to 50. There is also a levelling system, which upgrades player's character version (level) and gains memory for upgrades. Power Monger is a node control mode and Bit Runner is a capture-the-flag variation. Disintegration and team disintegration are classic deathmatch modes. Up to 10 people can play in online matches. Two modes focus on combat with light discs and two other modes feature vehicular combat on light cycles, as well as a special light tank vehicle. There are four different game modes in the game and four maps available initially. Evolution features persistent character progression system, that lets players earn new levels and unlock new upgrades, both in multiplayer and singleplayer. Each cycle leaves a light trail that can destroy or "derez" any enemy. The game also features light cycle races. Player's movements are heavily parkour-influenced, while combat was inspired by capoeira. The basic gameplay focuses on acrobatics and combat.
The game features both singleplayer and multiplayer modes. Tron: Evolution is a third-person action game with racing and role-playing game elements incorporated.